BSc(Hons) in Biology (Universitat de València, Spain, 2013), he completed his MSc in Paleontology (Universitat Autònoma de Barcelona, 2015) with a thesis entitled: New insights in a purported pterosaur pellet from the Upper Triassic of Friuli (NE Italy). From that time began his relation with ICP developed through the advisory of Josep Fortuny and Fabio M. Dalla Vecchia. Since 2016, Borja was granted by a PhD fellowship at the Museu Nacional / Universidade Federal do Rio de Janeiro (Brazil), where he is finishing his doctoral research under the advisory of Alexander W. A. Kellner and co-advisory of Josep Fortuny. His PhD focuses on the structure and evolution pterosaur postcranial skeletal pneumaticity compared with birds. His research also comprises other topics as with pterodactyloid phylogeny and pterosaur paleobiology, including the description of several new pterosaur species.